Situated in the centre of the site, Osterley Park House (listed grade I) consists of red-brick wings of three storeys around a raised courtyard with a taller brick tower with stone quoins and ogee cap projecting at each outer corner. The courtyard is entered through a grand Ionic portico on the east side. The portico stands at the head of a flight of steps and provides an entrance to the state rooms on the first floor, this being on a level with the courtyard. On the west side of the house a curved double staircase leads down from a pedimented door in the first-floor gallery to the garden. The mansion overlooks the east lawn with views to the Middle Lake.

The earliest mansion was built for Sir Thomas Gresham c 1575 and was largely transformed into its present design by the architect Robert Adam in the 1760s-1770s for the then owner Sir Francis Child. Although the original ground plan of Gresham's mansion was largely preserved the fabric was extensively altered, Adam continuing his work for Robert Child after the death of Sir Francis in 1763.

To the north of the mansion is the red-brick stable block (listed grade I). Built around a three-sided courtyard, the two-storey building has attics and turrets in the two internal angles. The open side of the courtyard faces the mansion. Now (2000) used to house a restaurant, shop, and other visitor facilities, the stables have their origins in the C16 and, with the exception of the clock tower and clock added in 1714, are little altered from the time of Thomas Gresham.
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